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(2015) discovered that exendin-4 treatment in Glp1r+/+ mice elevated cAMP levels in isolated intraepithelial lymphocytes while reducing the transcription of multiple inflammatory mediators, including IL-1, IL-6, IL-22, IL-12, TNF-, CCL2, CXCL1, and CXCL2

Yao C, Zhang F, Sun X, Shang D, He F, Li X, et al

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That is why its effect in studies extends far beyond the gastrointestinal tract, where it was originally discovered

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